Stamens of the Amaryllis with prominent anthers carrying pollen

Insects collecting nectar unintentionally transfer pollen to other flowers, causing pollination

The stamen (plural stamina or stamens, from Latin stamen meaning "thread of the warp") is the male organ of a flower. Each stamen generally has a stalk called the filament (from Latin filum, meaning "thread"), and, on top of the filament, an anther (from Ancient Greek anthera, feminine of antheros "flowery," from anthos "flower"), and pollen sacs, called microsporangia. The development of the microsporangia and the contained haploid gametophytes, (called pollen-grains) is closely comparable with that of the microsporangia in gymnosperms or heterosporous ferns. The pollen is set free by the opening (dehiscence) of the anther, generally by means of longitudinal slits, but sometimes by pores, as in the heath family (Ericaceae), or by valves, as in the barberry family (Berberidaceae). It is then dropped, or carried by some external agent — wind, water or some member of the animal kingdom — onto the receptive surface of the carpel of the same or another flower, which is thus pollinated.

Typical flowers have six stamens inside a perianth (the petals and sepals together), arranged in a whorl around the carpel (pistil). But in some species there are many more than six present in a flower (see, for example, the spider tree flower, below). Collectively, the stamens are called an androecium (from Greek andros oikia: man's house). They are positioned just below the gynoecium. The anthers are bilocular, i.e. they have two locules. Each locule contains a microsporangium. The tissue between the locules and the cells is called the connective.

In an immature, unopened flower bud, the filaments are still short. Their function is then to transport nutrients to the developing pollen. They start to lengthen once the bud opens. The anther can be attached to the filament in two ways:

• basifixed : attached at its base to the filament; this gives rise to a longitudinal dehiscence (opening along its length to release pollen)
• versatile : attached at its center to the filament; pollen is then released through pores (poricidal dehiscence).

Scanning electron microscope image of Penta lanceolata anthers, with pollen grains on surface

• monadelphous : fused into a single, compound structure
• diadelphous : joined partially into two androecial structures
• synantherous : only the anthers are connate (such as in the Asteraceae)

Stamens can also be adnate (fused or joined from more than one whorl):

• epipetalous : adnate to the corolla
• didynamous : occurring in two pairs of different length
• tetradynamos : occurring as a set of six filaments with two shorter ones
• exserted : extending beyond the corolla
• included : not extending from the corolla.

Plant sexuality

Stamen with pollinia and its anther cap. Phalaenopsis orchid.

In some species, however, the flowers are unisexual with only either male or female parts (monoecious = on the same plant; dioecious = on different plants). A flower with only male reproductive parts is called androecious. A flower with only female reproductive parts is called gynoecious.

A flower having only functional stamens is called a staminate flower.

An abortive or rudimentary stamen is called a staminodium, such as in Scrophularia nodosa.

The pistil and the stamens of orchids are fused into a column. The top part of the column is formed by the anther. This is covered by an anther cap.

Gallery

<gallery> Image:Crateva_religiosa.jpg|Flower of the spider tree (Crateva religiosa) with its numerous conspicuous stamens Image:WheatFlower3.JPG|Flowers of wheat at anthesis showing stamens. Image:Daylily Stamens dry 01v2.jpg|Stamens of a daylily (Hemerocallis), thickly covered in pollen Image:Albizia_julibrissin_'Rosea'_flower_detail.jpg|Flowers of the "silk tree" (Albizia julibrissin) have many long thread-like stamens Image:Passion flower stamen.jpg|Stamens of a passion flower (Passiflora caerulea) showing interesting symmetry </gallery>

References

ca:Estam cs:Ty?inka (botanika) de:Staubblatt es:Estambre eo:Stameno fa:???? (????) fr:Étamine io:Stamino id:Benang sari it:Stame la:Stamen lt:Kuokelis nl:Meeldraden ja:?? no:Pollenbærer pl:Pr?cik (botanika) pt:Estame ru:??????? fi:Hede sv:Ståndare tr:Erkek organ (çiçek) uk:??????? zh:??